Ok, thats ruled out then, bit like mine (hand made, point to point, 2 channel dual rectifer twin amp, based on fender '59 bassman clean channel and mesa/twin drive channel)
Is that a solid state rectifier cap where the GZ34 should be? Have you tried swapage there?
If you have swapped valves about, usually the phase inverter valve that makes the odd ocsalating sounds, and as it is a 'proper' amp, there is not a lot to go wrong!
I can give you the number of the guy wot builted mine but he is in Harrow.
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I remember when all this was fields..........